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Table 2 Partition of degrees of freedom for two loci with number of alleles n1 and n2 (a simplified case without distinguishing the paternal and maternal origins)

From: Models and partition of variance for quantitative trait loci with epistasis and linkage disequilibrium

Source Degrees of Freedom
additive (α) n1 - 1
dominance (δ) n1(n1 - 1)/2
additive (β) n2 - 1
dominance (γ) n2(n2 - 1)/2
additive × additive (αβ) (n1 - 1)(n2 - 1)
additive × dominance (αγ) (n1 - 1)n2(n2 - 1)/2
dominance × additive (δβ) n1(n1 - 1)(n2 - 1)/2
dominance × dominance (δγ) n1(n1 - 1)n2(n2 - l)/4
total n 1 ( n 1 + 1 ) 2 n 2 ( n 2 + 1 ) 2 1 MathType@MTEF@5@5@+=feaafiart1ev1aaatCvAUfKttLearuWrP9MDH5MBPbIqV92AaeXatLxBI9gBaebbnrfifHhDYfgasaacH8akY=wiFfYdH8Gipec8Eeeu0xXdbba9frFj0=OqFfea0dXdd9vqai=hGuQ8kuc9pgc9s8qqaq=dirpe0xb9q8qiLsFr0=vr0=vr0dc8meaabaqaciaacaGaaeqabaqabeGadaaakeaadaWcaaqaaiabd6gaUnaaBaaaleaacqaIXaqmaeqaaOGaeiikaGIaemOBa42aaSbaaSqaaiabigdaXaqabaGccqGHRaWkcqaIXaqmcqGGPaqkaeaacqaIYaGmaaGaeyyXIC9aaSaaaeaacqWGUbGBdaWgaaWcbaGaeGOmaidabeaakiabcIcaOiabd6gaUnaaBaaaleaacqaIYaGmaeqaaOGaey4kaSIaeGymaeJaeiykaKcabaGaeGOmaidaaiabgkHiTiabigdaXaaa@440F@