From: Holm multiple correction for large-scale gene-shape association mapping
AA
Aa
aa
MM
p 11 2
2 p 11 p 10
p 10 2
Mm
2 p 11 p 01
2 ( p 11 p 00 + p 10 p 01 )
2 p 10 p 00
mm
p 01 2
2 p 01 p 00
p 00 2